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"Partners of the night" are criminals Mary Regan, her uncle Joe Russell and Mathew Bradley, chief of detectives. Detective Clifford, assigned to the Mary Regan case, believes in her worth and so, rather than arresting Mary and her uncle for duping a bank teller into embezzling funds, Clifford forces them to make restitution. This angers Chief Bradley, who frames Clifford and then dismisses him from the force. However, Police Commissioner Thorne is in sympathy with the young detective and plots with him and Mary to crush Bradley. To frame the commissioner, Bradley instructs Mary to pass Thorne an envelope containing an alleged bribe, for which Mary substitutes an envelope containing blank paper, placing the bribe in the chief's cigar instead. When the reporters arrive, the chief is exposed, Clifford is reinstated, and he and Mary become partners for life.